Diastatidae

Diastatidae
	Diastata fuscula
	Diastatidae wing veins
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Superfamily:	Ephydroidea
Family:	Diastatidae; Hendel, 1917)
Type genus
	Diastata; Meigen, 1830
Subfamilies
	Diastatinae; Campichoetinae;

Diastata costata filmed in a forest near Marburg, Hesse; Germany.

Diastatidae are a family of flies in the order Diptera. They are encountered primarily in the Holarctic Region, but several species are found in the Oriental, Neotropical and Australasian regions. Members of the family number over 20 described species in three genera. There is an additional fossil genus.^[1]

Description

For terms see Morphology of Diptera

Minute flies with grey or brown-grey body and, usually, maculate wings. The postvertical bristles on head are cruciate and vibrissae are present on the head are present. The front orbital bristles are inset and upswept. The costa is interrupted near the end of Radial vein 1 and sometimes also near the humeral crossvein. The subcosta is incomplete fusing with Radial vein 1 before the apex. The posterior basal wing cell and discoidal wing cell are separate. The anal cell of wing and the anal vein of wing are both present.

Biology

Adults of living forms have been found along margins of bogs, marshes, and the edges of moist woodlands. Immature biologies are largely unknown. Hennig wrote about it thought to be Campichoeta punctum.

Genera

Diastatinae
- Diastata Meigen, 1830
Campichoetinae
- Campichoeta Macquart, 1835 (sometimes treated as a separate family Campichoetidae)
- Euthychaeta Loew, 1864
- †Pareuthychaeta Hennig 1965

Identification

Duda, O. (1934), Ephydridae. 6, 1, 58e, 1–18.In: Lindner, E. (Ed.). Die Fliegen der Paläarktischen Region 6: 1–115. Keys to Palaearctic species but now needs revision (in German).
A.A. Shtakel 'berg Family Diastatidae in Bei-Bienko, G. Ya, 1988 Keys to the insects of the European Part of the USSR Volume 5 (Diptera) Part 2 English edition. Keys to Palaearctic species but now needs revision .

Phylogeny

Ephydroidea

Curtonotidae

Drosophilidae+Camillidae

	Ephydridae

	Diastatidae sensu lato

Ephydroidea

	Curtonotidae+Drosophilidae

Campichoetidae

	Ephydridae+Camillidae

	Diastatidae

McAlpine (1989)^[2]

Grimaldi (1990)^[3]

Other

Diastatidae were once considered by some to be Ephydridae.

References

^ Papp, T. (1984). "Diastatidae". In Árpád Soós, Papp T. (ed.). Catalogue of Palaearctic Diptera (print). Vol. 10. Amsterdam: Elservier. pp. 182–185. ISBN 0-444-99601-X.
^ McAlpine, J.F. 1989. Chapter 116. Phylogeny and classification of the Muscomorpha. In Manual of Nearctic Diptera. Vol. 3. Coordinated by J.F. McAlpine and D.M. Wood. Agriculture Canada Monograph, 32. pp. 1397–1518.
^ David Grimald, 1990 A phylogenetic, revised classification of genera in the Drosophilidae (Diptera) Bulletin of American Museum of Natural History 1971-139 [1]

Wayne N. Mathis & David A. Barraclough Contributions to a Manual of Palaearctic Diptera. 3: 523–530. Science Herald, Budapest World Catalog and Conspectus on the Family Diastatidae (Diptera: Schizophora) Myia, 12:235–266 online here