Left: toes adducted (pulled towards the center) and spread (abducted); right, both feet clenched (plantar flexed)
The upper foot is clenching (plantarflexing) at the MTP joints and at the joints of the toes; the central foot is lifting the toes (dorsiflexing) at the MTP joints; and the foot flat on the ground off to the side is in a neutral position.
Kneeling with the MTP joints dorsiflexed (except the MTP joint of the little toe)